I think too many panels were too large. I know I sound like your typical "in my day" comic fan but you look back at some of the artistic masters from the past they use small panels to display action and on one page you got a whole sequence and that made it feel intense. Here you have a full page only showing a small part of a sequence. It looks pretty but to me, it's boring to read and doesn't really engage me as a reader.
One thing I've taken from my webcomics is to make every page count and get in as much as possible. Partly it's the typical webcomic updating (one page at a time so you want that page to count) and partly it's that when I tried the large panel after large panel people spent less time and were less engaged in reading it. And that's how I felt here. With large panel after large panel I just breezed through it. Not to mention that when you do try to keep the panel size down it's more jarring when you do go to the larger one, a trick you can use to shock the reader. Large panel after large panel just becomes boring.
This also makes me think of the iPod/iPad stuff. People talk about how when they read comics on them and go panel from panel it can adjust the size of the panel to fit the screen (enlarging small panels). I think that's wrong. Now you could say that artists have to compensate for it and make sure their panels fit well but I think it takes away some of the storytelling control the creators have. If you really want your comic to be viewable on the iPad in that way then perhaps you have to find some balance. To be honest, I have too many thoughts going on in my head and I could ramble about this one forever but I don't want to take over this rambling with another, I just wanted to put it out there.
Now I'm just an amateur webcomic guy so obviously I know less about making comics than your average (or pretty much any) professional. And this is all just my opinion, others are free (and apparently do) disagree. But it's how I feel.